Forum:Do the special damage bonuses apply to the upgraded damage or to the base damage?

Do the headshot damage/penetration/shield piercing/AP upgrades and/or cloak/adrenaline rush/etc bonuses apply to the upgraded damage or to the base damage of the weapon? For example, if I upgrade my weapon with all 5 damage upgrades, which gives +50% damage and then add an AP upgrade which gives me +50% damage to armor, would I get a total 200% or 225% of damage?